Central Question

Organelles are not isolated compartments. They exchange lipids and signals at membrane contact sites, generating local platforms that shape organelle identity, trafficking, metabolism, and stress responses.

The Ji Lab asks which molecular tethers and lipid transfer proteins assemble these sites, what cargo they move, and why their dysfunction contributes to neurological, metabolic, and developmental disease.

Schematic landscape of multiple organelles and contact sites
PDZD8 and Tex2 at ER-endolysosome contact sites
PDZD8 and Tex2 at ER-endolysosome contacts.

Project 01

ER-Endosome/Lysosome Contacts

ER-endolysosome contacts regulate organelle positioning, membrane identity, cargo sorting, and lysosomal function. The lab showed that PDZD8-mediated lipid transfer at ER-late endosome/lysosome contacts supports organelle positioning and neurite outgrowth.

Subsequent work identified Tex2 as a tubular ER protein recruited by TMEM55 to ER-late endosome/lysosome contacts, where it supports lysosomal trafficking, digestive capacity, and membrane lipid composition.

Gao et al., J Cell Sci, 2022 · Du et al., J Cell Biol, 2023

VPS13D and CHMP7 at ER-mitochondria and lipid droplet contacts
ER-mitochondria and lipid droplet contact biology.

Project 02

ER-Mitochondria and Lipid Droplet Contacts

ER-mitochondria contacts coordinate mitochondrial morphology, metabolism, and organelle quality control. The lab discovered that VPS13D interacts with VCP/p97 to negatively regulate ER-mitochondria contacts and mitochondrial organization.

Under starvation, VPS13D and TSG101 facilitate fatty acid transfer from lipid droplets to mitochondria. The lab also showed that CHMP7 can mediate three-way ER junctions and ER-mitochondria interactions outside its canonical ESCRT role.

Wang et al., Nat Commun, 2021 · Du et al., Mol Biol Cell, 2021 · Chu et al., Cell Death Differ, 2023

VPS13B and Sec23IP at the ER exit site-Golgi interface
VPS13B/COH1 and Sec23IP at ERES-Golgi interfaces.

Project 03

ER Exit Site-Golgi Interface and Cohen Syndrome

VPS13B/COH1 is the causative gene for Cohen syndrome, yet its molecular role had remained unclear. The lab identified Sec23IP as an adaptor that recruits VPS13B to ER exit site-Golgi interfaces and supports tubular ERGIC formation.

Disease-associated VPS13B missense variants disrupt Sec23IP binding, connecting a defined molecular interaction to defects in ER-to-Golgi transport and developmental disease biology.

Du et al., J Cell Biol, 2024

Sec14L6 transporting phosphoinositides between ER and lipid droplets
Sec14L6-dependent phosphoinositide transport at lipid droplets.

Project 04

Lipid Droplet Biogenesis

Lipid droplets form at the ER and require coordinated movement of phospholipids and signaling lipids. The lab identified Sec14L6 as a phosphoinositide transporter that regulates PI4P and PI(4,5)P2 homeostasis between lipid droplets and the ER.

Sec14L6 recruitment depends on ACSL3 at lipid droplets and PGRMC1 at the ER, providing a mechanistic view of lipid droplet biogenesis in mammalian cells.

Zhou et al., Nat Commun, 2025

SigmaR1 at ER sheets and the translocon
SigmaR1 links ER-sheet localization, lipid binding, and translocon function.

Project 05

ER Sheets and Translocon Control

ER sheets host ribosomes and the protein translocation machinery. The lab showed that SigmaR1 is an integral ER membrane protein enriched at ER sheets and functions as an auxiliary translocon factor with phosphatidylcholine-binding activity.

Loss of SigmaR1 disrupts protein and lipid droplet homeostasis, connecting ER-sheet organization to broad cellular physiology.

Hu et al., Nat Commun, 2026

Bridge-like lipid transfer protein at a membrane contact site
Bridge-like lipid transfer proteins at contact sites.

Project 06

Bridge-Like Lipid Transfer Proteins

Bridge-like lipid transfer proteins can span organelle interfaces and move lipids in bulk. The lab identified a BLTP1-SHIP164-BLTP2 axis that promotes Rab14-positive endosome bud formation at Golgi-endosome contacts.

This work links lipid transfer at membrane contact sites to membrane expansion, endosome budding, and retrograde trafficking.

Wang et al., Cell Discovery, 2024